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BCM32 Samenvatting College 5
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BCM32 College 5Cristy Verzijl
De glyoxylaat cyclus
Acetaat als koolstofbron in oa planten, gist en E.coli
Veel organismen, waaronder planten, kunnen acetaat gebruiken voor de biosynthese van
koolhydraten en voor de energievoorziening
Acetaat + CoA + ATP → acetyl CoA + AMP + PPi
PPi → 2 Pi
Deze organismen kunnen namelijk de glyoxylaatcyclus uitvoeren:
Acetaat komt als Acety‐CoA de glyoxylaatcyclus binnen en wordt halverwege de cyclus
omgezet in succinaat en carboxylaat:
Succinaat wordt in de citroenzuurcyclus verder verwerkt
Glyoxylaat wordt in de glyoxylaatcyclus verder verwerkt
Glyoxysomen
In planten vindt de glyoxylaatcyclus plaats in glyoxysomen.
Glyoxysomen ontwikkelen zich in lipide‐rijke zaden gedurende de ontkieming
Vetafbraak in plantenzaden
Plantenzaden kunnen door het uitvoeren van de glyoxylaatcyclus ook vetten gebruiken
voor de opbouw van koolhydraten, zoals glucose.
Via β‐oxidatie in de glyoxysomen:
Vetten → vetzuren → vet acylCoA → AcetylCoA → glyoxylaatcyclus → succinaat
De glyoxylaatcyclus omzeilt de 2 decarboxylatie stappen uit de citroenzuurcyclus om zo
vanuit een C2 molecuul een C4 molecuul te genereren.
Glyoxylaat cyclus in glyoxysoom
2
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