NADP+
ADP 3-Phosphoglycerate
LIGHT
+
REACTIONS:
Photosystem II Pi RuBP CALVIN
CYCLE
Electron transport chain
Photosystem I
ATP
Electron transport chain G3P
NADPH Starch
(storage)
Fotosysteem
Photosystem STROMA
Photon Light- Reaction-
harvesting center Primary
complexes complex electron
acceptor
Thylakoid membrane
Thylakoid membrane
Chlorophyll (green) STROMA
e–
Transfer Special pair of chloro-
of energy phyll a molecules Protein
Pigment
THYLAKOID SPACE molecules subunits THYLAKOID
(INTERIOR OF THYLAKOID) (purple) SPACE
(a) How a photosystem harvests light (b) Structure of a photosystem
2. Aangeslagen elektron v P680 è primaire elektronenacceptor (nu P680+)
3. H2O gesplitst dr enzymen è e- van de waterstofatomen naar P680+ è P680
a) P680+ is het sterkste bekende biologische oxidatiemiddel
b) De H+ komen vrij in de thylakoïde ruimte
c) O2 komt vrij als bijproduct van deze reactie
4. Elektronen è elektronentransportketen è PS I
(Vrijgekomen E zorgt voor creatie van protongradiënt over thylakoïdemembraan)
5. Epot opgeslagen in de protongradiënt stimuleert productie ATP door chemiosmose
6. In PS I (zoals PS II) maar P700
a) P700+ accepteert e- dat is doorgegeven van PS II via de
elektronentransportketen
7. Elektronen è NADP+-reductase
8. NADP+-reductase katalyseert overdracht e- & reduceert deze tot NADPH
a. e- van NADPH zijn beschikbaar voor reacties van de Calvincyclus
b. Dit proces verwijdert ook een H+ uit het stroma
Figure 11.16
Cyclische e- flow
Primary
• Alleen PS 1 gebruikt, e- komt op
Primary Fd
acceptor primaire acceptor en komt dan op
acceptor Fd
NADP+
cytochrome complex…
Pq
NADP+
reductase
+ H+
• Er wordt ATP gemaakt, geen O2/
Cytochrome
complex
NADPH
NADPH
Pc • Kan cellen beschermen tegen door
licht veroorzaakte schade
Photosystem I
Photosystem II ATP
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