STUDENT C O M P A N I O N
F U N D A M E N T A L S O F
Biochemistry
LIFE AT
THE MOLECULAR
LEVEL
Voet Voet Pratt
F O U R T H E D I T I O N
Akif Uzman Jerry Johnson Joseph Eichberg William Widger
Donald Voet Judith G. Voet Charlotte W. Pratt
,This page is intentionally left blank
rf rf rf rf rf
,STUDENT COMPANION TO ACCOMPANY rf rf rf
FUNDAMENTALS OF rf
BIOCHEMISTRY
LIFE AT THE MOLECULAR LEVEL
rf rf rf rf
Fourth Edition rf
Akif Uzman
rf
University of Houston rf rf
JerryJohnson rf
University of Houston rf rf
Joseph Eichberg rf
University of Houston rf rf
William Widger rf
University of Houston rf rf
Donald Voet rf
University of Pennsylvania rf rf
Judith G. Voet rf rf
Swarthmore College rf
Charlotte W. Pratt rf rf
Seattle, Washington rf
JOHN WILEY & SONS, INC
rf rf rf rf
, Cover Designer Madelyn Lesure Cov rf rf rf rf
er Illustration Norm ChristiansenCove
rf rf rf rf
r Photos
rf
(Vitruvian Man): © Odysseus/Alamy and © Dennis Hallinan/Alamy rf rf rf rf rf rf rf
Molecular structures clockwise from top: based on X- rf rf rf rf rf rf rf
ray structures that were respectively determined by Richard Dickerson and Horace Drew, Caltech; Gerard B
rf rf rf rf rf rf rf rf rf rf rf rf rf rf
unick, University of Tennessee; Thomas Steitz, Yale University; Alfonso Mondragón, Northwestern Unive
rf rf rf rf rf rf rf rf rf rf rf
rsity; Venki Ramakrishnan, MRC Laboratory of Molecular Biology, Cambridge, U.K.; Andrew Leslie and J
rf rf rf rf rf rf rf rf rf rf rf rf rf
ohn Walker, MRC Laboratoryof Molecular Biology, Cambridge, U.K.
rf rf rf rf rf rf rf rf
Founded in 1807, John Wiley & Sons, Inc. has been a valued source of knowledge and understanding for more t
rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf
han 200 years, helping people around the world meet their needs and fulfill their aspirations. Our company is b
rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf
uilt on a foundation of principles that include responsibility to the communities we serve andwhere we live and
rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf
work. In 2008, we launched a Corporate Citizenship Initiative, a global effort to address the environmental, soc
rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf
ial, economic, and ethical challenges we face in our business. Among the issues we are addressing are carbon i
rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf
mpact, paper specifications and procurement, ethical conduct within our business and among our vendors, and
rf rf rf rf rf rf rf rf rf rf rf rf rf rf r
community and charitable support. For more information, please visit our website: www.wiley.com/go/citize
f rf rf rf rf rf rf rf rf rf rf rf
nship.
Copyright © 2012 rf rf John Wiley & Sons, Inc. rf rf rf rf
All rights reserved. No part of this publication may be reproduced, stored in a retrieval system or transmitted i
rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf
n any form or by any means, electronic, mechanical, photocopying, recording, scanning or otherwise, except
rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf
as permitted under Sections 107 or 108 of the 1976 United States Copyright Act, without either the prior writt
rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf
en permission of the Publisher, or authorization through payment of the appropriate per-
rf rf rf rf rf rf rf rf rf rf rf rf
copy fee to the Copyright Clearance Center, Inc., 222 Rosewood Drive, Danvers, MA 01923, website www.c
rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf
opyright.com. Requests to the Publisher for permission should be addressed to theP ermissions Department, J rf rf rf rf rf rf rf rf rf rf rf rf rf rf
ohn Wiley & Sons, Inc. 111 River Street, Hoboken, NJ 07030-5774, (201) 748-6011, fax (201) 748-
rf rf rf rf rf rf rf rf rf rf rf rf rf rf rf
6008, website http://www.wiley.com/go/permissions.rf rf
To order books or for customer service, please call 1-800-CALL WILEY (225-5945).
rf rf rf rf rf rf rf rf rf rf rf
Student Companion (paperback) rf rf ISBN-13 978-1-1182-1827-3 r f
Printed in the United States of America1 rf rf rf rf rf rf rf
0 987 654 321
r f rf rf r f rf rf r f rf rf
